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Placement of exchange servers between AD sites

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We are planning to upgrade exchange 2013 and expand our exchange Organization. We have 10 AD sites with below details.

HQ Site 5000 Mailboxes

Site1 6000 Mailboxes

Site2 4000 Mailboxes

Site3 800 Mailboxes

Site4 500 Mailboxes

Site5-10  100-400 Mailboxes.

Below are our related concerns:

1.Do we need to place exchange servers in sites?  if yes, what are the requirements and criteria to place exchange server on sites.

if not, what are the bandwidth requirement in order to connect to HQ site.


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